The plant eaters take on the largely indigestible, spiny and poisonous defences of plants with some spectacular physical adaptations to diet. Plant Predators demonstrates the particular (and often peculiar) adaptations of herbivores

The third episode is devoted to the ways in which plants reproduce. Pollen and a stigma are the two components needed for fertilisation. Most plants carry both these within their flowers and rely on animals to transport the pollen from one to the stigma of another. To do this, they attract their couriers with colour, scent and nectar. It isn't just birds that help pollination: some mammals and reptiles also do so. However, it is mostly insects that are recruited to carry out the task. To ensure that pollen is not wasted by being delivered to the wrong flower, some species of plant have developed exclusive relationships with their visitors, and the gentian and its attendant carpenter bees is one example. Since pollen can be expensive to produce in terms of calories, some plants, such as orchids, ration it by means of pollinia and a strategically placed landing platform. Other orchids offer no reward for pollination, but instead mislead their guests by mimicking their markings and aroma, thus enticing males to 'mate' with them (Pseudocopulation). The most extreme fertilisation method is one of imprisonment, and one plant that uses it is the dead horse arum. It is often found near gull colonies, and mimics the appearance and smell of rotting flesh. Blow-flies are attracted to it, and are forced to stay the night before being allowed to depart in the morning, laden with pollen. Finally, Attenborough introduces the world's largest inflorescence: that of the titan arum.

Moving on to the Late Eocene period 36 million years ago and mammals have prospered and are now the largest creatures on land and sea. This is an era of animals like andrewsarchus, the biggest mammal carnivore ever to walk on land, and the brontotheres, small-brained herbivores. It is in the sea, however, that the most monstrous mammals of all can be found. We follow the fate of a female basilosaurus, a huge serpent-like early whale, but nothing like the gentle filter feeding whales of the 21st century. Four times the length of the great white shark, with jaws to match, she is every inch a killer.

25 million years ago the biggest land mammals of all time, the indricotheres, stalked the Earth. Up to seven metres tall and weighing 15 tonnes, adults were too big to be eaten by any predator of the time. Only in the first few years of its life was indricotheres vulnerable. The programme follows the fate of a calf from his traumatic birth to see whether it can survive droughts, killer hogs and hyaenodon - a predator the size of a rhino with jaws that could crush a rock.