The Cave of Swallows in Mexico is a 400m vertical shaft, deep enough to engulf the Empire State Building. The Lechuguilla cave system in the USA is 193km long and 500m deep with astonishing crystal formations hanging from its chambers. Although often overlooked, caves are remarkable habitats with equally bizarre wildlife. Cave angel fish cling to the walls behind cave waterfalls with microscopic hooks on their flattened fins. Cave swiftlets navigate by echo-location and build nests out of saliva. The Texas cave salamander has neither eyes nor pigment. Unique access to a hidden world of stalactites, stalagmites, snotites and troglodytes brings a wealth of surprises.

Dr Nigel Spivey explores how art influences life by tracing the development of the image from cave paintings to our modern obsession with images. Dr. Spivey begins his investigation by travelling to the Cave of Altamira near the town of Santillana del Mar in Cantabria, Spain, where in 1879 a young girls exclamation of 'Papa, look, oxen!' to her father, local amateur archaeologist Marcelino Sanz de Sautuola, is explained to have meant that Maria had just become the first modern human to set eyes on the first gallery of prehistoric paintings ever to be discovered.

Open your eyes to the bizarre, ferocious and surprisingly beautiful world of the invertebrates, a ground-breaking exploration into a spectacular miniature universe never normally seen but teeming all around us. In the first episode, the story of the land-living invertebrates. Discover the private life of Europe's dramatic leopard slug, a common garden resident with a truly bizarre end to its marathon mating ritual; watches the courtship ballet of tiny springtails on the underside of a leaf; sees swarms of bright red South African millipedes find partners, and in the caves of Venezuela meets the giant bat-eating centipede.

It's 3.5 million years ago and in East Africa a remarkable species of ape roams the land. Australopithecus afarensis has taken the first tentative steps towards humanity by standing and walking on two legs. The rift valley was forming, and the rain forests dying as Africa dried out - turning the landscape into a mosaic of scattered trees and grass. In this new environment afarensis found it more efficient to move about on two legs rather than four.

'Blood Brothers' follows the lives of two species, Paranthropus boisei and Homo habilis who embody two alternative ways of ape-man life. Although heavyset, with distinctive gorilla-like faces, the boisei are gentle characters. They live within a strict social structure and are led by a dominant male whose strength and power holds the group together. The habilis have taken a different approach to survival. They don't have the specialisms of the boisei but instead have developed into the archetypal jack-of-all-trades, inquisitive scavengers prepared to try almost anything to survive. Tough, active, gregarious and noisy, they are always on the move and always alert to the possibility of a meal. But in the near drought of the dry season the habilis are struggling. It seems as if their way of life cannot help them when conditions are tough.

One and a half million years ago, a new breed of ape-man walks the land. In southern Africa, Homo Ergaster has taken the next step to becoming human. They have long, modern looking noses, which cool air as they breathe. Their hairless bodies, with millions of tiny sweat glands, mean they don't pant anymore to control their temperature - they sweat. And, above all, they have big brains - nearly two-thirds the size of ours.