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East Africa is a land which is constantly changing. To survive here, creatures must be able to deal with unpredictable twists and turns - wet turning to dry, feast to famine, cold to hot - no matter how hostile it becomes. From dense forests to snow capped peaks, steamy swamps and endless savannah, this unique and varied land is also a haven for life, supporting large animals in numbers found nowhere else on Earth. But away from the familiar, forever-travelling herds, there are a huge cast of other characters - lizards that steal flies from the faces of lions, vast dinosaur-like birds who stalk catfish through huge wetlands, and an eagle who risks everything on the arrival of ten million bats from a far off rainforest.

Mammals dominate the planet. They do it through having warm blood and by the care they lavish on their young. Weeks of filming in the bitter Antarctic winter reveal how a mother Weddell seal wears her teeth down keeping open a hole in the ice so she can catch fish for her pup. A powered hot air balloon produces stunning images of millions of migrating bats as they converge on fruiting trees in Zambia, and slow-motion cameras reveal how a mother rufous sengi exhausts a chasing lizard. A gyroscopically stabilised camera moves alongside migrating caribou, and a diving team swim among the planet's biggest fight as male humpback whales battle for a female.

About 340 million years ago a brand new family of animals was evolving in the primeval swamps. They were to go one step further than the amphibians who had emerged onto dry and before them. For they would eventually completely cut their ties with water. They were the ancestors of todays lizards. They evolved scaly impermeable skins and moved up into the forests. They diversified into a multitude of different shapes and sizes. They developed signalling systems to communicate with one another. And they squabbled as animals do. For food they hunted insects that were already well established on the land in great numbers. And here without returning to water they produced their families. They powered their bodies not only with food but with the heat that they drew directly from the sun.
As they diversified so they spread into the harshest of the lands habitats. The baking waterless deserts which eventually they would come to dominate. Discover jacky lizards that wave, wrestling beaded lizards and the the world's smallest chameleon, which is no bigger than his thumbnail, and the biggest lizard in Australia.

In a late Jurassic forest in what is now China, an Epidexipteryx escapes from a juvenile Sinraptor by climbing a tree. It finds a beetle grub in the tree bark, being shown to use its elongated fingers in a similar way to a modern day aye-aye. However, its prey is stolen by another, larger Epidexipteryx, and after a brief bout of posturing, the smaller individual goes to find more food. It drops a second grub to the forest floor, where the other Epidexipteryx retrieves it, only to be killed by the juvenile Sinraptor. The episode then cuts to a desert in late Cretaceous Mongolia, where a Saurornithoides is shown brooding a nest of eggs. When it leaves the nest, an Oviraptor raids it, fleeing when the troodontid returns. The Saurornithoides is suddenly attacked and eaten by a Gigantoraptor, which then heads to compete in a breeding ritual for mates. The males use their feathers for display, a brief fight between two erupting at one point, allowing the females to choose the best suitor. The episode finally cuts to an early Cretaceous forest in China, where a Xianglong is being hunted by a Microraptor, which uses its feathers to pursue the gliding lizard in the air. A Sinornithosaurus attacks it, and after a brief chase the Microraptor manages a lucky escape. The Sinornithosaurus, alongside two other members of its species is then shown hunting a Jeholosaurus and its three young. The group brings down the parent, the narrator explaining that their possibly venomous bite allowed them to tackle animals much larger than themselves. A montage is then shown of the feathered dinosaurs featured in the programme, with the narrator saying that Microraptor not only hints at how flight might have developed, but also that dinosaurs still live amongst us today, as birds.

No two islands in the Galapagos are the same. The imperceptible drift of a continental plate keeps each island biologically isolated. David Attenborough explores this evolutionary crucible, encountering tortoises that weigh up to half a tonne, finches that use tools and lizards that communicate using press-ups; for Darwin, this was all evidence for his theory of evolution. We see the final footage of the world famous tortoise fondly known as Lonesome George, the last survivor of his species. David Attenborough was the last person to have ever filmed with him. Darwin’s famous visit had a downside – the arrival of man. David investigates the impact we’ve had in these islands, as our influence is a double-edged sword. We’ve disrupted the natural balance but he also believes Darwin would be thrilled with the advances we have made in science. We’re also now uncovering evidence that evolution is more rapid than Darwin could ever have imagined. Whatever wonders the Galapagos Islands hold today, they are only a hint of what awaits them in the future.

Will Smith discovers hidden worlds of fast and slow in the Earth's oldest desert. Worlds where things go so fast or so slow we don't even know they're happening. This hidden world of speed is everywhere, even 80 feet underwater. Hundreds of tiny anemones, anchored to the rocks, but film them patiently, then speed it up, and this miniature world comes to life. A lizard's tongue is one of the fastest movements in the entire animal kingdom. An orca creates a shock wave by slapping her tail and that pulse can travel through the water at more than a thousand miles an hour. to stuns the herrings she hunts.