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Northern Africa is home to the greatest desert on Earth, the Sahara. On the fringes, huge zebras battle over dwindling resources and naked mole rats avoid the heat by living a bizarre underground existence. Within the desert, where the sand dunes 'sing', camels seek out water with the help of their herders and tiny swallows navigate across thousands of square miles to find a solitary oasis. This is a story of an apocalypse and how, when nature is overrun, some are forced to flee, some endure, but a few seize the opportunity to establish a new order.

The second programme deals with the mechanics of flight. Getting into the air is by far the most exhausting of a bird's activities, and Sir Attenborough observes shearwaters in Japan that have taken to climbing trees to give them a good jumping-off point. The albatross is so large that it can only launch itself after a run-up to create a flow of air over its wings. A combination of aerodynamics and upward air currents (or thermals), together with the act of flapping or gliding is what keeps a bird aloft. Landing requires less energy but a greater degree of skill, particularly for a big bird, such as a swan. Weight is kept to a minimum by having a beak made of keratin instead of bone, a light frame, and a coat of feathers, which is maintained fastidiously. The peregrine falcon holds the record for being fastest in the air, diving at speeds of over 300 km/h. Conversely, the barn owl owes its predatory success to flying slowly, while the kestrel spots its quarry by hovering. However, the true specialists in this regard are the hummingbirds, whose wings beat at the rate of 25 times a second. The habits of migratory birds are explored. After stocking up with food during the brief summer of the north, such species will set off on huge journeys southwards. Some, such as snow geese, travel continuously, using both the stars and the sun for navigation. They are contrasted with hawks and vultures, which glide overland on warm air, and therefore have to stop overnight.

The next instalment focuses on dietary needs and how different species have evolved beaks to suit their individual requirements. The latter come in a multitude of forms. Blue tits and goldfinches have beaks akin to tweezers, with which to extract seeds, while the hawfinch's razor-like bill can deal with a cherry-stone. However, the crossbill is the only finch that can twist its mandibles in opposite directions. Jays store acorns for winter by burying them in the ground, whereas woodpeckers can keep up to 60,000 of them in one tree trunk. Sap is also desirable, and there are a variety of methods used to obtain it. The hoatzin is the only specialised leaf-eater, and accordingly has a digestive system more akin to that of cattle. Plants recruit birds to aid pollination, and offer nectar as a reward. Hummingbirds eat little else, and the sword-bill's beak is the longest of any bird in relation to its body. Insects are also highly prized, and Galapagos finches are shown to possess some ingenuity as they not only strip bark, but also use 'tools' to reach their prey. Crows are hailed as being among the most intelligent birds, and one is shown using a twig to spear a grub within a fallen log. The robin is an opportunist, and Sir Attenborough observes one seizing morsels as he digs a patch of earth. In South America, a cattle tyrant sits atop an obliging capybara and uses its vantage point to spot passing food that may be dislodged by its grazing partner.

The fifth programme explores the alliances formed between the animal and plant worlds. Attenborough dives into Australia's Great Barrier Reef and contrasts the nocturnal feeding of coral, on microscopic creatures, with its daytime diet of algae. Some acacias are protected by ants, which will defend their refuge from any predator. Besides accommodation, the guards are rewarded with nectar and, from certain species, protein for their larvae as well. Fungi feed on plants but can also provide essential nutriment to saplings (Mycorrhiza). The connection is never broken throughout a tree's life and a quarter of the sugars and starches produced in its leaves is channelled back to its fungal partners. Meanwhile, fungi that feed on dead wood leave a hollow trunk, which also benefits the tree. Orchids enjoy a similar affiliation. Lichens are the product of a relationship between fungi and a photosynthetic associate, usually algae. They are extremely slow-growing, and a graveyard is the perfect location to discover their exact longevity. Mistletoe is a hemiparasite that obtains its moisture from a host tree, while using own leaves to manufacture food. Its seeds are deposited on another by the mistletoe tyrannulet, following digestion of the fruit. The dodder (Cuscuta) is also parasitic, generally favouring nettles, and siphons its nourishment through periodic 'plugs' along its stem. The rafflesia has no stem or leaves and only emerges from its host in order to bloom — and it produces the largest single flower: one metre across.

Once life arrived in the Galapagos, it exploded into unique and spectacular forms. David Attenborough investigates the driving forces behind such evolutionary innovations. We learn that life must be able to adapt quickly in these ever-changing volcanic landscapes. It has resulted in species found nowhere else in the world, such as giant whale sharks and marine iguanas that can spit sea-salt from their noses, dandelion seeds that grow into tree-sized plants and spiders that can blend perfectly into the darkness. Adaptation has been the key to survival in these islands so far, but the story of life in the Galapagos doesn’t end here. The catalyst that triggers these explosions of life remains in place.

No two islands in the Galapagos are the same. The imperceptible drift of a continental plate keeps each island biologically isolated. David Attenborough explores this evolutionary crucible, encountering tortoises that weigh up to half a tonne, finches that use tools and lizards that communicate using press-ups; for Darwin, this was all evidence for his theory of evolution. We see the final footage of the world famous tortoise fondly known as Lonesome George, the last survivor of his species. David Attenborough was the last person to have ever filmed with him. Darwin’s famous visit had a downside – the arrival of man. David investigates the impact we’ve had in these islands, as our influence is a double-edged sword. We’ve disrupted the natural balance but he also believes Darwin would be thrilled with the advances we have made in science. We’re also now uncovering evidence that evolution is more rapid than Darwin could ever have imagined. Whatever wonders the Galapagos Islands hold today, they are only a hint of what awaits them in the future.