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On the coast, two worlds collide. Coasts are the most dynamic and challenging habitats in the ocean - that brings great rewards but also great danger. The extraordinary animals that live here must find ingenious ways to cope with two very different worlds. This episode is a rollercoaster ride of heart-stopping action and epic drama, peopled with characters from the beautiful to the bizarre. We meet fish that live on dry land and puffins that must travel 60 miles or more for a single meal, and witness a life-and-death struggle in a technicolour rock pool. In a secluded cove in the Galapagos, sea lions feast on 60kg tuna. It should be impossible - tuna are usually far too fast for sea lions to catch. But here the sea lions club together to herd their prey inshore. Once trapped in the shallows, these huge fish are easy pickings. As the tide recedes in Brazil, lightfoot crabs leap from rock to rock, desperately avoiding the water - their lives depend on it. Moray eels launch themselves from rock pools, jaw gaping. Then octopuses, too. Both crawl across dry rock to set their ambush. Elsewhere, the ever-changing tides create rock pools. But these temporary worlds are a battleground. Predatory starfish turn a magical garden into the stuff of nightmares.

The story of the British Library's Royal Manuscripts collection reaches its end with the last great flowering of illumination, in the magnificent courts of the Tudors. Dr Janina Ramirez investigates astrological texts created for Henry VII, and unwraps his will - still in its original, extravagantly-decorated velvet and gold cover. She hears music written for Henry VIII, which went unperformed for centuries; and reads love notes between the king and Anne Boleyn, written in the margins of a prayer book. Nina also visits Bruges, the source of many of the greatest manuscripts, where this medieval art form collided with the artistic innovations of the Renaissance.

Sultan Mehmed II's siege of Constantinople enters its third week. Mehmed's artillery teams continue their relentless bombardment of Constantinople's walls. After his navy suffers a devastating defeat, Mehmed moves his ships overland to the Golden Horn in a daring, visionary feat. In the shadow of betrayal, Giustiniani attacks the Ottoman fleet.

Exoplanets, strange worlds outside our solar system. We're discovering the cosmos is full of alien planets. Alien worlds that challenge our understanding of planetary systems. Hellishly hot worlds, violently colliding worlds, worlds getting eaten by their stars. There's much, much more out there than we had ever imagined.
Exoplanets are shaking up our understanding of the universe. The cosmos is a chaotic array of the odd, the weird, and the wonderful. The more we find, the less we know.

Immense shoals of fish throng our shallow seas. Small fish, in turn, sustain bigger ones. The rich coastal seas are the fishing grounds of our planet and can provide an abundance of food for wildlife and humanity. The seas fringing land make up less than a tenth of the world's oceans but 90 percent of marine creatures live in coastal waters, from fearsome sharks to lowly urchins. Protecting these habitats is a battle humanity must win.

Fueled by a ruthless and maniacal ambition to conquer Central Japan, Nobunaga has destroyed many enemies who have underestimated him. His vicious military campaigns to weaken the powerful Buddhist institutions have turned large swathes of the population against him. As Nobunaga's cruelty intensifies, some generals begin to question his command, leading to a betrayal that alters the political landscape forever.