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A humpback whale mother and calf embark on an epic journey from tropical coral paradises to storm ravaged polar seas. Newly discovered coral reefs in Indonesia reveal head-butting pygmy seahorses, flashing 'electric' clams and bands of sea kraits, 30-strong, which hunt in packs. Elsewhere plagues of sea urchins fell forests of giant kelp. Huge bull fur seals attack king penguins, who despite their weight disadvantage, put up a spirited defence.

There is no greater test for life than winter, as temperatures plummet to 70 below and winds reach 200kph. Darkness and ice extend across the polar regions and only a few remarkable survivors gamble on remaining. We join a female polar bear trekking into the Arctic mountains to give birth as the first blizzards arrive. Out on the frozen ocean, the entire world's population of spectacled eider ducks brave the winter in a giant ice hole kept open by ferocious currents. Arctic forests transform into a wonderland of frost and snow - the scene of a desperate and bloody battle between wolf and bison, but also where a remarkable alliance between raven and wolverine is made. Beneath the snow lies a magical world of winter survivors. Here tiny voles dodge the clutches of the great grey owl, but cannot escape the ultimate under-show predator - the least weasel. Midwinter and a male polar bear wanders alone across the dark, empty icescape. Below the snow, polar bear cubs begin life in an icy den while fantastical auroras light the night skies above. In Antarctica, we join male emperor penguins in their darkest hour, battling to protect precious eggs from fierce polar storms. Weddell seals escape to a hidden world of jewel-coloured corals and alien-looking creatures but frozen devastation follows as sinister ice stalactites reach down with deadly effect.

The fifth programme explores the alliances formed between the animal and plant worlds. Attenborough dives into Australia's Great Barrier Reef and contrasts the nocturnal feeding of coral, on microscopic creatures, with its daytime diet of algae. Some acacias are protected by ants, which will defend their refuge from any predator. Besides accommodation, the guards are rewarded with nectar and, from certain species, protein for their larvae as well. Fungi feed on plants but can also provide essential nutriment to saplings (Mycorrhiza). The connection is never broken throughout a tree's life and a quarter of the sugars and starches produced in its leaves is channelled back to its fungal partners. Meanwhile, fungi that feed on dead wood leave a hollow trunk, which also benefits the tree. Orchids enjoy a similar affiliation. Lichens are the product of a relationship between fungi and a photosynthetic associate, usually algae. They are extremely slow-growing, and a graveyard is the perfect location to discover their exact longevity. Mistletoe is a hemiparasite that obtains its moisture from a host tree, while using own leaves to manufacture food. Its seeds are deposited on another by the mistletoe tyrannulet, following digestion of the fruit. The dodder (Cuscuta) is also parasitic, generally favouring nettles, and siphons its nourishment through periodic 'plugs' along its stem. The rafflesia has no stem or leaves and only emerges from its host in order to bloom — and it produces the largest single flower: one metre across.

The deep is perhaps the most hostile environment on earth, at least to us - a world of crushing pressure, brutal cold and utter darkness. We have barely begun to explore it, and yet it is the largest living space on the planet. Scientists already think that there is more life in the deep than anywhere else on earth. This episode takes us on an epic journey into the unknown, a realm that feels almost like science fiction. We discover alien worlds, bizarre creatures and extraordinary new behaviours never seen before. We encounter savage hordes of Humboldt squid hunting lanternfish in the depths and coral gardens flourishing in absolute darkness, with more species of coral to be found in the deep than on shallow tropical reefs. Narrated by David Attenborough.

Corals build themselves homes of limestone in the warm, clear, shallow seas of the tropics. Their reefs occupy less than one tenth of one per cent of the ocean floor, yet they are home to a quarter of all known marine species. The broad-club cuttlefish has found its place by using a hypnotic display that apparently mesmerizes its prey, causing it to let down its defences. On the Great Barrier Reef a remarkable grouper uses sign language, dubbed the headstand signal, to reach out to an entirely different creature, a reef octopus, to flush small fish out of their hiding holes and into the groupers waiting mouth.

Far from land, where few of us ever venture, is the ocean beyond the boundary of any country. Largely ungoverned, wild, and lawless. Venture into the deep, dark and desolate oceans that are home to an abundance of beautiful - and downright strange - creatures.