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Rome was once a largely democratic society, with regular elections. This Republic lasted for 500 years, but then came Tiberius Gracchus. He believed in the ideals of the Republic - fairness, decency and justice for everyone -but was appalled by Rome's aristocrats' treatment of the poor. So he unleashed the power of the mob upon the streets of Rome, with devastating consequences.

In the final part, Professor Al-Khalili uncovers tales of success and heartache in the story of chemists' battle to control and combine the elements, and build our modern world. He reveals the dramatic breakthroughs which harnessed their might to release almost unimaginable power, and he journeys to the centre of modern day alchemy, where scientists are attempting to command the extreme forces of nature and create brand new elements.

Fish dominate the planet's waters through their astonishing variety of shape and behaviour. The beautiful weedy sea dragon looks like a creature from a fairytale, and the male protects their eggs by carrying them on his tail for months. The sarcastic fringehead, meanwhile, appears to turn its head inside out when it fights. Slow-motion cameras show the flying fish gliding through the air like a flock of birds and capture the world's fastest swimmer, the sailfish, plucking sardines from a shoal at 70 mph. And the tiny Hawaiian goby undertakes one of nature's most daunting journeys, climbing a massive waterfall to find safe pools for breeding.

Birds owe their global success to feathers - something no other animal has. They allow birds to do extraordinary things. For the first time, a slow-motion camera captures the unique flight of the marvellous spatuletail hummingbird as he flashes long, iridescent tail feathers in the gloomy undergrowth. Aerial photography takes us into the sky with an Ethiopian lammergeier dropping bones to smash them into edible-sized bits. Thousands of pink flamingoes promenade in one of nature's greatest spectacles. The sage grouse rubs his feathers against his chest in a comic display to make popping noises that attract females. The Vogelkop bowerbird makes up for his dull colour by building an intricate structure and decorating it with colourful beetles and snails.

Fresh water is our most precious resource and it defines the distribution of life on land. Follow the descent of rivers from their mountain sources to the sea. Watch spectacular waterfalls, fly inside the Grand Canyon and explore the wildlife below the ice in the world's deepest lake. Witness unique and dramatic moments of animal behaviour: a showdown between smooth-coated otters and mugger crocodiles; deep-diving long tailed macaques; massive flocks of snow geese on the wing and a piranha frenzy in the perilous waters of the world's largest wetland.

The Cave of Swallows in Mexico is a 400m vertical shaft, deep enough to engulf the Empire State Building. The Lechuguilla cave system in the USA is 193km long and 500m deep with astonishing crystal formations hanging from its chambers. Although often overlooked, caves are remarkable habitats with equally bizarre wildlife. Cave angel fish cling to the walls behind cave waterfalls with microscopic hooks on their flattened fins. Cave swiftlets navigate by echo-location and build nests out of saliva. The Texas cave salamander has neither eyes nor pigment. Unique access to a hidden world of stalactites, stalagmites, snotites and troglodytes brings a wealth of surprises.