A jaguar fights to keep his river home in Brazil as the night brings both challenges and opportunities to hunt. Evening is falling in the world's greatest tropical wetland, Brazil's Pantanal. A stronghold for South America's most secretive big cat, the jaguar. And enjoying the last warmth of the day is a male known as Juru. At six years old and weighing over 100 kilos, he's amongst the largest, most powerful big cats in the Americas. This prime stretch of river has been his territory for two years. But tonight Juru's reign hangs in the balance. In a recent fight with a rival male, he badly injured his paw. It's left him weak and hungry. The coming days and nights will test him more than ever before. He'll need to summon the strength to defend his river.

Deep in the twisted roots of a giant strangler fig tree, hidden from view, lives perhaps the most bizarre nighttime creature of all. The spectral tarsier, tiny primates barely larger than a tennis ball. As darkness falls, a family of these small primates in Sulawesi ventures into the jungle to find food while protecting their babies.

In the fourth episode of the series, legendary nature and cultural photographer Art Wolfe captures the great brown bear with the stunning backdrop of Alaska’s mountains and glaciers, immigrating wildebeests in the plains East Africa, and the hard-to-find gorillas of Uganda.

In the fourth episode, leading landscape photographer Peter Eastway follows in the footsteps of Frank Hurley – the pioneering Australian photographer on Shackleton’s expedition to the south a century ago – to capture the wilderness and wildlife of Antarctica and South Georgia.

Once life arrived in the Galapagos, it exploded into unique and spectacular forms. David Attenborough investigates the driving forces behind such evolutionary innovations. We learn that life must be able to adapt quickly in these ever-changing volcanic landscapes. It has resulted in species found nowhere else in the world, such as giant whale sharks and marine iguanas that can spit sea-salt from their noses, dandelion seeds that grow into tree-sized plants and spiders that can blend perfectly into the darkness. Adaptation has been the key to survival in these islands so far, but the story of life in the Galapagos doesn’t end here. The catalyst that triggers these explosions of life remains in place.

No two islands in the Galapagos are the same. The imperceptible drift of a continental plate keeps each island biologically isolated. David Attenborough explores this evolutionary crucible, encountering tortoises that weigh up to half a tonne, finches that use tools and lizards that communicate using press-ups; for Darwin, this was all evidence for his theory of evolution. We see the final footage of the world famous tortoise fondly known as Lonesome George, the last survivor of his species. David Attenborough was the last person to have ever filmed with him. Darwin’s famous visit had a downside – the arrival of man. David investigates the impact we’ve had in these islands, as our influence is a double-edged sword. We’ve disrupted the natural balance but he also believes Darwin would be thrilled with the advances we have made in science. We’re also now uncovering evidence that evolution is more rapid than Darwin could ever have imagined. Whatever wonders the Galapagos Islands hold today, they are only a hint of what awaits them in the future.