The open arenas of grassland and desert make up half of all land on our planet. In these exposed habitats, predators like cheetahs, bald eagles and lions can usually see their prey. But it works both ways: their prey can see them too. With nothing but open vistas, the element of surprise is hard-won, and predators must make their own opportunities.

A jaguar fights to keep his river home in Brazil as the night brings both challenges and opportunities to hunt. Evening is falling in the world's greatest tropical wetland, Brazil's Pantanal. A stronghold for South America's most secretive big cat, the jaguar. And enjoying the last warmth of the day is a male known as Juru. At six years old and weighing over 100 kilos, he's amongst the largest, most powerful big cats in the Americas. This prime stretch of river has been his territory for two years. But tonight Juru's reign hangs in the balance. In a recent fight with a rival male, he badly injured his paw. It's left him weak and hungry. The coming days and nights will test him more than ever before. He'll need to summon the strength to defend his river.

Deep in the twisted roots of a giant strangler fig tree, hidden from view, lives perhaps the most bizarre nighttime creature of all. The spectral tarsier, tiny primates barely larger than a tennis ball. As darkness falls, a family of these small primates in Sulawesi ventures into the jungle to find food while protecting their babies.

In the fourth episode of the series, legendary nature and cultural photographer Art Wolfe captures the great brown bear with the stunning backdrop of Alaska’s mountains and glaciers, immigrating wildebeests in the plains East Africa, and the hard-to-find gorillas of Uganda.

In the fourth episode, leading landscape photographer Peter Eastway follows in the footsteps of Frank Hurley – the pioneering Australian photographer on Shackleton’s expedition to the south a century ago – to capture the wilderness and wildlife of Antarctica and South Georgia.

Once life arrived in the Galapagos, it exploded into unique and spectacular forms. David Attenborough investigates the driving forces behind such evolutionary innovations. We learn that life must be able to adapt quickly in these ever-changing volcanic landscapes. It has resulted in species found nowhere else in the world, such as giant whale sharks and marine iguanas that can spit sea-salt from their noses, dandelion seeds that grow into tree-sized plants and spiders that can blend perfectly into the darkness. Adaptation has been the key to survival in these islands so far, but the story of life in the Galapagos doesn’t end here. The catalyst that triggers these explosions of life remains in place.