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Grasslands cover one quarter of all land and support the vast gatherings of wildlife, but to survive here animals must endure the most hostile seasonal changes on the planet. From Asia's bizarre-looking Saiga antelope to the giant anteaters of Brazil, grassland animals have adapted in extraordinary ways to cope with these extremes. In the flooded Okavango, lions take on formidable buffalo in epic battles, on the savannah bee-eaters take advantage of elephants to help catch insects and, on the freezing northern tundra, caribou embark on great migrations shadowed by hungry Arctic wolves.

Cities are growing at a faster rate than any other habitat on Earth. They may seem an unlikely place for animals to thrive, but they can be a world of surprising opportunity. Leopards prowl the streets of Mumbai, peregrine falcons hunt amongst New York's skyscrapers, and a million starlings perform spectacular aerial dances over Rome. In Jodhpur, langurs are revered as religious deities and in Harar, locals live in harmony with wild hyenas. Many animals, however, struggle to cope in the urban jungle. As the architects of this environment, can humans choose to build cities that are homes for both them and wildlife?

In 1914, Russia entered World War I in response to Austria-Hungary's declaration of war on Russia's ally Serbia. This was fought across multiple fronts while isolated from its allies. The Battle of Tannenberg, where the entire Russian Second Army was annihilated, cast an ominous shadow over the empire's future. The already-existing public distrust of the regime was deepened by the rising costs of the war and rumours of corruption. World War I takes Nicholas to the front. Back home, outrage grows over high casualties and Alexandra and Rasputin's political intrigues.

In the third episode, the discovery of extraterrestrial life might face an impossible challenge: the physics of the universe itself; but using cutting-edge tech, experts might be on the verge of a groundbreaking find -- and the evidence could already be in hand.

After ending 120 years of civil war and reunifying Japan under his banner, Toyotomi Hideyoshi, the supreme ruler of Japan, lays dead. In a desperate attempt to protect his dynasty, just before his death, Hideyoshi created a council of five powerful warlords. They are to govern Japan until his young son comes of age. But the power hungry Tokugawa Ieasyu declares war on those who oppose him.

The Caribbean is home to a notorious band of British pirates called the Flying Gang. They plunder merchant ships, sailing from the Americas to Europe and Africa. England's King George I has had enough, so he comes up with a shock tactic to end piracy. It's an offer from the King, a pardon, clemency for those who turn their backs on piracy. Those who refuse will be hunted down and hanged. Two pirates, Blackbeard and Charles Vane stand the King's man, Woodes Rogers. Who will win the final battle?