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Diarmaid MacCulloch's own life story makes him a symbol of a distinctive feature about Western Christianity - scepticism, a tendency to doubt which has transformed both Western culture and Christianity. In the final programme in the series, he asks where that change came from. He challenges the simplistic notion that faith in Christianity has steadily ebbed away before the relentless advance of science, reason and progress, and shows instead how the tide of faith perversely flows back in. Despite the attacks of Newton, Voltaire, the French Revolutionaries and Darwin, Christianity has shown a remarkable resilience. The greatest damage to Christianity was actually inflicted to its moral credibility by the two great wars of the 20th century and by its entanglement with Fascism and Nazism. And yet it is during crisis that the Church has rediscovered deep and enduring truths about itself, which may even be a clue to its future.

Travelling forward in time to 30,000 years ago, it's the middle of an ice age. The landscape is dominated by mighty mammoths, living side-by-side with woolly rhinos, giant deer and two separate species of human. The programme follows the fate of a herd of mammoths in their annual struggle against the harsh ice-age conditions. Every summer they spend on the grassy plains of what will one day become the bottom of the North Sea, but every winter they are forced to head for the less exposed valleys further south. It is a journey fraught with danger: mammoths can get trapped in frozen bogs, and the herd must run the gauntlet of hunters like cave lions and the deadly Neanderthals.

In the far south west of Japan, there is a chain of islands stretching towards the tropics - a place where all life is influenced by the power of the sea, and where volcanoes and typhoons are forces to be reckoned with. The journey begins at an island at the top of the chain and travels south, revealing unexpected stories of isolation, unique wildlife and unsolved mysteries.

Nathan and Angela accelerate their parenting rehearsal with three-year-old, then six-year-old actors portraying their fake son, 'Adam'. Angela refuses to participate in Halloween due to her belief in Satanic conspiracies.
Nathan balances parenting duties with a rehearsal for Patrick, a man who wants to confront his brother over their late grandfather's will, which bans Patrick from inheriting money if he is dating a 'gold digger'. The rehearsal occurs in a replica Raising Cane's restaurant in a warehouse next to the relocated Alligator Lounge. To introduce real emotions to the rehearsal, Nathan stages a scenario to convince Patrick that he could inherit buried gold from the grandfather of Isaac, the actor who plays Patrick's brother. After Patrick has an emotional breakthrough during a rehearsal, he leaves the production and never returns. Nathan narrates that he is envious that self-deception is easy for some people as he ponders his fake family.

Corals build themselves homes of limestone in the warm, clear, shallow seas of the tropics. Their reefs occupy less than one tenth of one per cent of the ocean floor, yet they are home to a quarter of all known marine species. The broad-club cuttlefish has found its place by using a hypnotic display that apparently mesmerizes its prey, causing it to let down its defences. On the Great Barrier Reef a remarkable grouper uses sign language, dubbed the headstand signal, to reach out to an entirely different creature, a reef octopus, to flush small fish out of their hiding holes and into the groupers waiting mouth.

The big blue is the world's greatest wilderness, far from shore and many kilometres deep. It's a vast marine desert where there is little to eat and nowhere to hide. Yet it's home to some of the biggest and most spectacular creatures on earth. This episode reveals what it takes to survive in this savage and forbidding world. We witness feats of incredible endurance, moments of high drama and extraordinary acts of heart-wrenching self-sacrifice. Every animal in the big blue must find their own unique way to survive. Sperm whales have the largest brains in the world. They live for 80 years, and we are only now beginning to learn the extraordinary complexity of their language of clicks - thought to coordinate the whole family in everything from childcare to hunting. With special pressure-proof cameras, we witness record-breaking feats of endurance as they hunt for squid a kilometre down into the abyss. Only recently have we begun to solve the mystery of where baby turtles disappear to in their early years. They leave the crowded waters of the coast and head to the open ocean, where they use floating debris like logs as life rafts. Here they remain until adulthood, adrift on the high seas in relative safety away from coastal predators. Over half of all animals in the open ocean drift in currents. Jellyfish cross entire oceans feeding on whatever happens to tangle with their tentacles. The jelly-like Portuguese man-of-war can harness sail power to fish with its deadly tentacles. Sometimes there is a brief explosion of food in this marine desert, but ocean hunters must be fast to make the best of this bonanza. We witness super pods of up to 5,000 spinner dolphins racing to herd vast shoals of lanternfish, briefly caught at the surface where it is thought they spawn. New aerial footage reveals, for the first time, the truth to a centuries-old sailors' legend of the 'boiling seas' - the spectacular feeding frenzy of 90kg tuna and dolphins smashing through the lantern fish shoals turning the sea white with foam.